<p>In 1858, Charles Darwin was forty-nine years old, a gentleman scientist living quietly at Down House in the Kent countryside. He was not yet a focus of debate; his "big book on species" still lay on his desk as a manuscript. For more than twenty years he had been accumulating material for it, puzzling over the questions that it raised, trying to bring it to a satisfactory conclusion, and wanting to be certain that his startling theory of evolution was correct. </p><p>It is at this point that the concluding volume of Janet Browne's magisterial biography opens. Beginning with the extraordinary events that finally forced the <i>Origin of Species</i> into print, we come to the years of fame and controversy. Here, Browne does dramatic justice to all aspects of the Darwinian revolution, from a fascinating examination of the Victorian publishing scene to a survey of the debates between scientists and churchmen over evolutionary theory. At the same time, she presents a wonderfully sympathetic and authoritative picture of Darwin himself.</p>